SAT Question 5fd86f4b: Form Structure And Sense answer and explanation

English - Form Structure And Sense - Subject Verb Agreement
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The ghazal, a poetic form originating in seventh-century Arabic poetry, has an intricate structure. The twentieth-century Kashmiri American poet Agha Shahid Ali explains that each one of a ghazal’s couplets, while adhering to the patterns of rhyme (qafia) and refrain (radif) established in the poem’s opening lines (matla), ______ thematically and logically autonomous, resulting in a poem with “a stringently formal disunity.”
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Difficulty:Hard
Which choice completes the text so that it conforms to the conventions of Standard English?