SAT Question d10e46a2: Transitions answer and explanation

English - Transitions - Example Or Illustration
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The ancient Spartans were known for their bitingly concise—or laconic—wit, a quality they maintained even in the face of great peril. ______ when Philip II of Macedon threatened Laconia (the region of Greece containing Sparta), he said, “If I invade Laconia, I shall turn you out.” The Spartans replied with a single word: “If.”
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Difficulty:Easy
Which choice completes the text with the most logical transition?