SAT Question db2e480a: Form Structure And Sense answer and explanation

English - Form Structure And Sense - Verb Tense Consistency
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By the time Hawaiian king Kamehameha III ______ the throne, the number of longhorn cattle, first introduced to the islands in 1793, had drastically increased, and so too had the need for paniolo (Hawaiian cowboys) to manage the wild herds that then roamed throughout the volcanic terrain.
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Difficulty:Easy
Which choice completes the text so that it conforms to the conventions of Standard English?