SAT Question e2759b92: Form Structure And Sense answer and explanation

English - Form Structure And Sense - Verb Tense Consistency
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Occupying a significant part of modern-day Nigeria, the Kingdom of Benin was one of the major powers in West Africa between the thirteenth and nineteenth centuries. It ______ ruled by Oba Ewuare I from 1440 to 1473.
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Difficulty:Easy
Which choice completes the text so that it conforms to the conventions of Standard English?