SAT Question f0864217: Form Structure And Sense answer and explanation

English - Form Structure And Sense - Subject Modifier Placement
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Rabinal Achí is a precolonial Maya dance drama performed annually in Rabinal, a town in the Guatemalan highlands. Based on events that occurred when Rabinal was a city-state ruled by a king, ______ had once been an ally of the king but was later captured while leading an invading force against him.
195
Difficulty:Hard
Which choice completes the text so that it conforms to the conventions of Standard English?