SAT Question 39ac6498: Boundaries answer and explanation

English - Boundaries - End Of Sentence Punctuation
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In forecasting weather events, meteorologists sometimes discuss the role of atmospheric rivers. What are atmospheric rivers, and how ______ Part of the water cycle, atmospheric rivers are narrow channels of moisture moving through the atmosphere. In certain conditions, these “rivers” can release some of their moisture as precipitation.
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Difficulty:Easy
Which choice completes the text so that it conforms to the conventions of Standard English?