SAT Question 6181924a: Boundaries answer and explanation

English - Boundaries - No Punctuation Within Clause
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Fernando Palma Rodríguez creates robotic sculptures that combine mechanical elements with materials like feathers, soil, and seeds. The artist is from a rural farming community outside Mexico City, and he studied engineering in college. The natural and mechanical ______ highlight these two aspects of his background.
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Difficulty:Hard
Which choice completes the text so that it conforms to the conventions of Standard English?