SAT Question 69f031ab: Form Structure And Sense answer and explanation

English - Form Structure And Sense - Finite And Nonfinite Verb Forms
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While exploring Nevada’s Gypsum Cave in 1930, Seneca and Abenaki archaeologist Bertha Parker made her most famous discovery: the skull of a now-extinct ground sloth (Nothrotheriops shastensis) alongside human-made tools. Parker’s crucial finding was the first ______ humans in North America as far back as 10,000 years ago.
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Difficulty:Medium
Which choice completes the text so that it conforms to the conventions of Standard English?