SAT Question 9902d2de: Boundaries answer and explanation

English - Boundaries - Comma And Coordinating Conjunction Between Clauses
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The Alvarez theory, developed in 1980 by physicist Luis Walter Alvarez and his geologist son Walter Alvarez, maintained that the secondary effects of an asteroid impact caused many dinosaurs and other animals to die ______ it left unexplored the question of whether unrelated volcanic activity might have also contributed to the mass extinctions.
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Difficulty:Easy
Which choice completes the text so that it conforms to the conventions of Standard English?