SAT Question aa443c4b: Form Structure And Sense answer and explanation

English - Form Structure And Sense - Finite And Nonfinite Verb Forms
0:00
FREE OFFICIAL BANK
Recent pollen analyses of the Aran Islands have led some researchers to propose that the now treeless islands were once wooded. This hypothesis ______ that certain trees, such as P. sylvestris, survived without interruption or human intervention throughout the Holocene cannot stand, researchers Michael O’Connell and Karen Molloy counter, unless other explanations can first be ruled out.
136
Difficulty:Hard
Which choice completes the text so that it conforms to the conventions of Standard English?