SAT Question b0fb36ad: Form Structure And Sense answer and explanation

English - Form Structure And Sense - Subject Verb Agreement
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Unique among animal species, humans use our vocal apparatuses primarily for two separate communicative purposes: to talk and to sing. The question of what cross-cultural traits distinguish these distinct modes, and secondarily what pressures led humans to develop them in the first place, ______ neuropsychologist Daniela Sammler’s 2024 study “Signatures of Speech and Song: ‘Universal’ Links despite Cultural Diversity.”
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Difficulty:Hard
Which choice completes the text so that it conforms to the conventions of Standard English?