SAT Question cd2443c0: Form Structure And Sense answer and explanation

English - Form Structure And Sense - Subject Modifier Placement
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A pioneer in the field of taphonomy (the study of how organic remains become fossils), ______ may be just as prevalent in the fossil record as those of thick-shelled organisms.
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Difficulty:Medium
Which choice completes the text so that it conforms to the conventions of Standard English?