SAT Question 0205e563: Transitions answer and explanation

English - Transitions - Contrast Transition
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At two weeks old, the time their critical socialization period begins, wolves can smell but cannot yet see or hear. Domesticated dogs, ______ can see, hear, and smell by the end of two weeks. This relative lack of sensory input may help explain why wolves behave so differently around humans than dogs do: from a very young age, wolves are more wary and less exploratory.
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Difficulty:Medium
Which choice completes the text with the most logical transition?