SAT Question 05a14e18: Boundaries answer and explanation

English - Boundaries - Punctuating Items In A Series
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While the ancient Celts—Iron Age peoples who inhabited parts of western and central Europe—weren’t a single unified group, their art often featured common elements. These included intricate patterns of interlocking spiral lines, which often held symbolic ______ of birds, horses, and other animals; and inlaid enamel accents.
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Difficulty:Hard
Which choice completes the text so that it conforms to the conventions of Standard English?