SAT Question 267a13e2: Boundaries answer and explanation

English - Boundaries - Period Between Sentences
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In 2010, archaeologist Noel Hidalgo Tan was visiting the twelfth-century temple of Angkor Wat in Cambodia when he noticed markings of red paint on the temple ______ the help of digital imaging techniques, he discovered the markings to be part of an elaborate mural containing over 200 paintings.
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Difficulty:Medium
Which choice completes the text so that it conforms to the conventions of Standard English?