SAT Question 89ab0d46: Boundaries answer and explanation

English - Boundaries - No Punctuation Within Clause
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After the printing press was introduced in 1440, handwritten manuscripts from Europe’s medieval period were often destroyed and the paper used for other purposes. In one instance, pages ______ a collection of Norse tales dating to 1270 were discovered lining a bishop’s miter (hat).
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Difficulty:Easy
Which choice completes the text so that it conforms to the conventions of Standard English?