SAT Question f114cbf0: Transitions answer and explanation

English - Transitions - Contrast Transition
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A firefly uses specialized muscles to draw oxygen into its lower abdomen through narrow tubes, triggering a chemical reaction whereby the oxygen combines with chemicals in the firefly’s abdomen to produce a glow. ______ when the firefly stops drawing in oxygen, the reaction—and the glow—cease.
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Difficulty:Medium
Which choice completes the text with the most logical transition?